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How To Visit The Temple Of Poseidon At Cape Sounion, Athens

The Temple of Poseidon at Sounion is an easy half-day trip from Athens. Here’s everything you need to know about planning a Cape Sounion sunset tour in 2024.

Temple of Poseidon Athens Greece

The Temple of Poseidon at Cape Sounion

The Temple of Poseidon at Sounio is one of the most majestic temples in the whole of Greece. It is located at the southernmost tip of the Attica peninsula, about 70 kms (43.5 miles) out of central Athens.

No Greek God or Goddess could have asked for a better spot to have a temple built in their honor. Cape Sounio is surrounded by steep rocks, offering spectacular views to the Aegean Sea. 

In my opinion, it’s one of the most beautiful places to visit in Greece .

Sunset at the temple of Poseidon at Sounion

Many people prefer to visit the ancient site during the evening, when the sun sets behind the temple and illuminates it in a beautiful orange hue. Watching the sun set from the top of the cliff is an unforgettable experience.

There are three ways to visit the Poseidon temple: by public transportation, rental car, or an organized tour / taxi / private transfer.

Take the KTEL bus to the temple of Poseidon at Sounion

Public buses to Sounio depart from a bus stop opposite Pedion tou Areos , close to Victoria metro station on the green line. (A guide here on how to use the metro in Athens ).

These buses are known as KTEL buses, and they take around 2 hours to get to the temple, or longer during peak hours. On summer weekends, when Athenians go out to the beach, there will generally be more traffic. A one way ticket costs about 6.5 euro.

Cape Sounion Temple of Poseidon

The bus timetables are seasonally adjusted, and there are fewer routes during the off-season. You can find bus timetables  here , though unfortunately the website is not regularly updated. In fact, your best best would be to call and ask.

If you visit Cape Sounio by bus, you will need to make sure that you don’t miss the last bus to return to Athens, as there are very few hotels in the area.

Rent a car and drive to Cape Sounio

Another way to get to the temple of Poseidon at Sounion is to hire a car. This way, you will have more flexibility and you won’t worry about the return bus back to Athens, especially if you want to stay for sunset.

Check out an online platform called Discover Cars to compare prices and car models, and book a rental car to drive to Sounio.

The coastal route on the way to Sounio, known as the Athens Riviera, is very scenic, and you will enjoy lovely views of the Aegean. There are many hidden coves and sandy beaches along the way, where you can stop for a swim.

Visit the temple of Poseidon at Sounion

It will take you about an hour and a half to get to the temple from the Athens centre. Free parking is available outside the site.

You can then go for lunch / dinner to the nearby port town of Lavrio, known for its wonderful fish tavernas. There’s also the – rather indifferent – Akrogiali restaurant on the small beach nearby, close to hotel Aegeon Beach .

Driving in Greece

If you are thinking to drive to Sounio, you should know that the coastal road to Sounio has many twists and turns, and is narrow at times.

View of the Athens riviera

If you are coming back after dark, pay extra attention, as long parts of the road are poorly lit. Or you could take the inland route, which is not very picturesque but it’s much smoother to drive.

One more thing to consider is that Greeks are pretty fast drivers. In my experience having driven this road dozens of times, there are always people speeding or driving in the middle of the road.

This article on driving in Greece will give you an idea of what to expect.

Take an organized Cape Sounion Sunset Tour

Another option is to take a half-day tour to Sounion. There are numerous companies offering this service, as a group tour or a private tour.

Sometimes you will have a tour guide who will show you around the Poseidon temple. Other companies will leave you to explore the site at your own pace. Read the tour descriptions carefully!

If you want to catch the sunset from the temple of Poseidon, make sure you choose one of the evening tours.

Here are a few tour options:

  • Sunset tour to temple of Poseidon
  • Guided tour to the temple of Poseidon (morning)
  • Guided tour to the temple of Poseidon (evening)
  • Fun kayak trip to the temple of Poseidon – worth splashing out for!

Visiting the Temple of Poseidon, Sounion

Once you have arrived at the parking area, make sure you park the car safely. There are some toilets here if  you need them, and note that there aren’t any inside the ancient site. There’s also a cafe where you can have a coffee or drink.

You can then enter the site, and buy your ticket at the ticket booth. For the time being, this is the only way to get your ticket. 

You will then go up the hill to get to the imposing temple. The path is not flat, but it’s not too steep – nothing like the path leading up to the Acropolis Hill.

The temple is built on the highest point of the hill. Out of the 34 original marble columns, 15 are still standing. Some people might spot the name “Byron”, which is carved on one of the columns, a sign that Lord Byron was here.

View of the temple of Poseidon at Cape Sounio

The views to the sea are really out of this world, and in my opinion this is what makes the temple of Poseidon so special. I’m sure the God of the Sea was very happy to have his temple built on this spot!

There are various trails and paths leading down to the side of the cliff. It’s worth following one or two of them and spending some time looking out to the sea, especially when it’s close to sunset.

Make sure you bring water, sunscreen, a hat, comfy shoes, and your camera! 

Tickets and opening hours

Temple of Poseidon tickets cost 10 euro from April to October, and 5 euro from November to March, and you can only get them at the ticket booth. Entrance is free on certain days:

  • Melina Mercouri Day (6 March)
  • World Heritage Day (18 April)
  • International Museums Day (18 May)
  • European Heritage Days (last weekend of September)
  • OXI day (28 October) 
  • Every first Sunday in November, December, January, February and March.

For more information, have a look at this guide on how to visit the ancient sites in Greece for free .

The Temple of Poseidon opening hours are from 9:30 to sunset, which is when many people try to visit. As sunset times vary throughout the year, make sure you check in advance.

And now that you know how to get to this wonderful temple and what to see, let’s have a look at some facts!

Early history of Cape Sounio

According to archaeological evidence, the wider area of Cape Sounion had been considered sacred since the Bronze Age.

Homer mentions Sounio in his famous poem, the Odyssey. It was one of the places where the Greek fleet stopped on their return from Troy.

A statue of Poseidon or Zeus at the National Archaeological Museum in Athens

The temple of Poseidon is also the place where King Aegeus, the father of the hero Theseus, committed suicide jumping from the cliff, thinking his son had been killed by the Minotaur.

As a tribute to King Aegeus, his name was given to the Aegean Sea, where most of the Greek islands are located. Here’s an introduction to the Greek island groups .

The Temple of Poseidon replaced an earlier temple

The existing Poseidon Temple was built between 444-440 BC, on top of the ruins of an Archaic temple. The white marble used for its construction came from the nearby town of Lavrion, which was best known for its rich silver mines. Remains of the older temple, dating from the archaic period, are not visible.

Poseidon temple Athens half day trip

A couple of decades later, during the Peloponnesian war, the Cape was enclosed by an extensive wall, which was considered the strongest existing fortification in the proximity of Athens. Remains of these defensive walls still survive today.

Cape Sounion was a strategic spot for the city-state of ancient Athens. Any boat sailing towards Piraeus would pass by the cape. Moreover, the rich mines at Lavrion enabled Athens to achieve unprecedented wealth and power during the 5th century BC.

The Sea God Poseidon

So who was Poseidon, and why was he so important in Ancient Greece?

Poseidon was one of the Gods of Mount Olympus , and the son of the Titans Cronus and Rhea. He is always depicted as a serious-looking, bearded man holding his powerful weapon, the trident.

Poseidon Ancient Greek God of the Sea

Among Greeks, Poseidon was known as the God of the Sea, and was responsible for the fates of sailors. He also controlled fresh waters, rivers, pools and lakes, and regulated earthquakes and natural disasters.

Finally, Poseidon was a fan of horses and horse races. His chariot was pulled by imaginary monstrous creatures, half-horses and half-snakes, while dolphins, fish and numerous other sea creatures followed along.

People worshiped the mighty Poseidon all around the Ancient Greek world. Many of his temples survive, not only close to the coast, but also up on the mountains, close to lakes and rivers.

A few trivia about God Poseidon

Like many other gods, Poseidon was particularly fertile. According to Greek mythology, he fathered over 100 children, with several goddesses and mortal women. At the same time, he was also attracted by men, such as Niritis and Pelopas.

Poseidon’s legitimate wife was Queen Amphitrite, with whom he had a son, Triton, and by some accounts a daughter, Benthesikyme. Pegasus, the flying horse, was the offspring of Poseidon and Medusa.

Fame riding Pegasus

Unlike the children of Zeus, some of Poseidon’s descendants were evil and malicious. Allegedly, Poseidon himself killed some of his villainous sons, while others were eliminated by Hercules and Theseus.

There are hundreds of myths, stories and legends connected with the God of the sea.

An Athens temple for Athena and Poseidon

A famous story has to do with the myth of Athena and Poseidon, or how the city-state of Athens took its name .

The Parthenon in Athens

The two Gods competed over the powerful city state of Athens, and Athena won. The Athenians built the Parthenon, the most famous temple in the ancient site of the Acropolis, in her honor.

Poseidon was furious, and he used his superpowers to flood an area outside Athens, the Thriasian Plain. To calm him down, the Athenians built another temple, called Erechtheion, close to the Parthenon. This was a temple dedicated to both Athena and Poseidon.

Interestingly, no earthquakes have ever caused any damage to the Acropolis. This can be attributed to the fact that Ancient Greeks were excellent engineers. Or, maybe, the God of Earthquakes protected the ancient site, because his own temple was there. I’ll leave it to you to decide!

The Caryatids statues in the Erechtheion temple dedicated to Athena and Poseidon

FAQ about the Temple of Poseidon at Sounio

Here are a few questions that visitors often ask:

Where is Poseidon’s temple located?

The location of Poseidon’s temple is about 70 kms (43.5 miles) south of central Athens.

How many columns did the Temple of Poseidon have?

In ancient times, the temple of Poseidon had 34 columns made out of white marble. Out of those, fifteen survive today. On one column, you can spot the name of Lord Byron, the famous English poet, who visited the ancient temple in 1810.

Is the Temple of Poseidon worth seeing?

Definitely! The Poseidon Temple is one of the most beautiful ancient sites in Greece. If you’re visiting Athens, it’s absolutely  worth making a trip down to Sounion to see the temple up close, and enjoy the fantastic sea views.

How do you get to the Temple of Poseidon?

There are three ways to get to the Temple of Poseidon in Sounion. You can take a KTEL bus, rent a car, or take a tour / private taxi.

Can you swim at the Temple of Poseidon?

The Temple of Poseidon is built up on a hill. If you have your own transportation, there are many beaches nearby where you can go for a swim.

God Poseidon and his trident

More Athens travel guides

If you enjoyed this article, have a look at these other ones:

  • How to spend 3 days in Athens
  • When is the best time to go to Athens?
  • Plaka neighborhood in Athens
  • The 20 best museums in Athens
  • 15 Free things to do in Athens Greece
  • How to get from Athens to Thessaloniki by train

Vanessa from Real Greek Experiences

Hi! I’m Vanessa from Athens. The ancient site at Cape Sounion is one of my favourites in Greece, not just because of the temple, but also because of the incredible views. If you have a few days in Athens, make sure to visit. Follow my FB page for more Greece-related info! 

4 thoughts on “How To Visit The Temple Of Poseidon At Cape Sounion, Athens”

Awesome post, so very informative and detailed. I will be reaching Athens in June at 1.30pm and probably will take an hour to leave my bags. Will be staying for 3 nights. However, I am confused, how can I cover on my Day 1 if I follow your Day 2 and Day 3 as is?

Hi! I’m assuming you are referring to this post ? I guess you could drop one or two activities from Days 2 or 3, I’m not sure you could fit everything in, but it all depends on how fast you like doing things, and what you are mostly interested in!

Tudo bem? Como faço para comprar somente os ingressos para o Templo de Poseidon? Procuro na internet e só acho venda de passeios guiados. Estarei de carro e só quero realmente comprar os ingressos. Existe algum site oficial que só vende os ingressos? Agradeço se puder me orientar. Obrigada, Adriana

Oi! You will just need to buy the tickets at the entrance, at the moment there is no other way!

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Half-Day Sounion Day Trip from Athens with the Temple of Poseidon

poseidon temple tour

  • Embark on an extraordinary journey to the Temple of Poseidon at Sounion, located just outside Athens, where you'll witness the timeless majesty of the temple and the breathtaking sunset views.
  • Let an expert archaeologist guide you through the ancient mysteries and captivating stories that surround this magnificent site, immersing you in a thrilling exploration of Greece's rich history and mythical tales.

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Inspiring Sounio: Temple of Poseidon Sunset Trip

Cape Sounio, Athens

Celebrate the magic of the Temple of Poseidon with an Athens to Cape Sounio fully guided trip led by an expert, state-licensed guide from our team. As you journey towards the realm of Poseidon, explore the hidden treasures of the 'Athens Riviera' and uncover local havens for sun and sea. Immerse yourself in the timeless grandeur of the ancient Temple of Poseidon and witness the breathtaking spectacle of the sun setting over the Aegean Sea, an experience that transcends time and legend.

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HIGHLIGHTS WHAT YOU CAN EXPECT PRACTICAL INFO

Temple of Poseidon in Cape Sounion

The meeting point for this half-day guided excursion from Athens to Cape Sounio is the lobby of your hotel.

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Inclusions and Exclusions

  • The Location
  • Important Information

Know before you book

Know before you go, cape sounion: the temple of poseidon.

poseidon temple tour

€  7,99 per person

Contributors and Bibliography

Everyone knows the beauty of the sunsets in Cape Sounion. Far fewer people, though, realise that this is one of Athens’ most impressive archaeological sites, dedicated to a god known for his moody approach to life and home to a vibrant community of soldiers, priests, and slaves that turned ancient Athens into a superpower.

Waltz in Cape Sounion with a self-guided audio tour on your phone designed by an accredited expert. Listen to fascinating stories, based on an award-winning storytelling concept, and gain great insights into the most iconic landmark of Cape Sounion, the temple of Poseidon. Start your tour from the entrance of the cape and continue to meet the temple, getting acquainted with its surroundings, and the temple of Athena Sounias.

This is an unmissable opportunity to experience at your own pace the Cape Sounion with a captivating audio tour on your phone.

* * Ticket/entrance fee to the Temple of Poseidon is not included.

The entrance to the archaeological site is to the east of the sanctuary, a few meters from the intercity bus stop, the souvenir shop and the cafe. There is limited parking here, so it is best to leave the car in the lower parking lot. The tour’s first stop is on the plateau with the information panel dedicated to “The Fortress: the Wall” to the right of the trail just after the ticket office.

poseidon temple tour

Leonidas Argyros

  • Self-guided audio tour in English on your smartphone (Android & iOS)
  • An activation link to access your audio tour
  • Offline content (text, audio narration and maps) to avoid roaming charges
  • Entrance tickets to the Temple of Poseidon are not included.
  • Smartphone or headphones
  • Food and drinks
  • Transportation
  • App not compatible with Windows phones
  • Hotel pick-up and drop-off

Mandatory items

  • Charged smartphone

The location

Starting point.

There is no meeting point. The self-guided audio tour is designed to start at the Temple of Poseidon entrance (Cape Sounio, Sounio 195 00).

poseidon temple tour

Directions to starting point

The easiest way to get there is by reaching the Mavromateon Bus stop (Athens 104 34) which will take you to Cape Sounion. At the bus station, there will be a sign plate with the Cape Sounion name on it.

  • North side. The temple of Poseidon
  • East facade. Byron’s graffiti’
  • The Sanctuary of Athena

Important information

Recommended visiting hours.

Before sunset

Additional admission

November 1st – March 31st: 5€

April 1st – October 31st: 10€

  • Please consider that admission fees for Cape Sounion are not included.
  • This is a downloadable self-guided audio tour for your smartphone, accessible through our app. No live guide will accompany you. No VR / AR is included.
  • No live guide or meeting point provided.
  • After booking, you will receive an email with further instructions on how to access and download your audio tour. Check your spam folder as well.
  • Book per device to be used, not per participant.
  • An Android (version 5.0 and later) or iOS (version 15.0 and later) smartphone is required. The audio tour is not compatible with Windows Phones, iPhone 6/6 Plus or older, iPod Touch 6th generation or older, iPad 4th generation or older, iPad Air 1st generation, iPad Mini 3rd generation or older.
  • You will need storage space on your phone (100-150 MB).
  • Mobile signal may be weak at the site(s). Download your audio tour on your smartphone prior to your visit to fully enjoy it.
  • After booking, you will receive an email with further instructions on how to access and download your audio tour. Please check your email spam folder as well
  • Ensure that your smartphone is fully charged & that you bring your earphones
  • Visitors are advised to wear comfortable shoes 

Contributors

Image Credits:

Central: © Dimitris Polydoropoulos / Commons Wikimedia

Stop 1: © Carole Raddato / Commons Wikimedia

Stop 2: © Ava Balili / Flickr

Stop 3: © Tumblr

Stop 4: © George E. Koronaios / Commons Wikimedia

Stop 5: © Nikos Zacharioudakis

Stop 6: © Nikos Zacharioudakis

Stop 7: © Mirjanamimi / Commons Wikimedia

Download the app

To take this self-guided tour, you will need to download the Clio Muse Tours app on your iOS or Android device. Enjoy your tour!

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Cape Sounion and Temple of Poseidon Tickets

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Cape Sounion and Temple of Poseidon | Day tour from Athens

The Temple of Poseidon is an imposing monument perched atop a hill, dedicated to the formidable sea god Poseidon. Unlike Percy Jackson, ancient Greeks could not get away so easily by disobeying Olympian gods! Hence, they built this temple to seek favors for their journeys. The drive from Athens to Cape Sounion is about 90 minutes and offers picturesque views of the Athenian coastline. You can also catch splendid views of the sunset along the Aegean Sea from Poseidon’s Temple. Here’s a comprehensive guide to planning your visit to the Temple of Poseidon at Cape Sounion.

Why visit the Cape Sounion and Temple of Poseidon?

Why visit the Cape Sounion and Temple of Poseidon?

Scenic day tour from Athens

The biggest highlight of visiting the Temple of Poseidon is the scenic views of the Saronic Gulf, from Athens to Cape Sounion, along the Athinon Souniou Road. Whether you travel by car or public transport, you can catch glimpses of many of the iconic UNESCO Heritage sites in and around Athens like the Temple of Zeus, the Acropolis, the archaeological site of Thorikos, Glyfada Beach, and more. You can take detours and explore these sites up close or continue your journey towards Cape Sounion. 

Sunset views from Cape Sounion

Don’t miss the sunset from Cape Sounion. The Temple of Poseidon provides a unique vantage point from where you can watch the sun set upon the Aegean Sea. The rays illuminate the temple’s marble columns, offering stellar photo opportunities. On clear days, you can see the Saronic Gulf islands from a distance. As the day ends, the Temple of Poseidon’s location offers a calming backdrop to contemplate and relax. 

Marvel at Doric architecture

Like other temples built during the Classical period, the Temple of Poseidon also featured Doric columns, harmonious proportions, and religious inscriptions. However, its columns are much thinner and more refined than other temples built around that era. The sleek columns allowed for more light to enter the site and made it seem higher than usual. Poseidon's Temple was said to have around 34 huge columns, out of which only around 15 are standing today.

Offers a glimpse into Greek mythology

Cape Sounion is a haven for history buffs and lovers of mythology. In Homer’s Odyssey, King Aegeus, the famous king of Athens, plummets to his death from Sounion. It is said that the group of sailors who were returning to Athens did not put away their black sails. King Aegeus spotted their ships and believed his son, Theseus, had passed away in battle. Unable to handle his grief, the king took his life. Visit the temple site and walk around the ruins, from where this incident is said to have taken place. 

Explore nearby ruins and beaches

If you ever played Assassin's Creed: Odyssey, you would know that the Temple of Poseidon is not just a stand-alone ruin. Just like the videogame, there are several side quests to explore in the Cape Sounion archaeological site. You can visit a smaller Temple of Athena Sounias nearby, or head to the adjacent Sounio Beach. If you have a car at your disposal, visit Lake Vouliagmeni, around 20 km away from Sounion. Swim in the mineral-rich waters of this lake and enjoy a serene escape from your daily worries.

Things to know before booking your Temple of Poseidon tickets

  • Leave behind your transport woes as you plan your visit to Poseidon’s Temple. All Cape Sounion tours include round-trip transfers to and from Athens in luxurious, air-conditioned vehicles. 
  • There is a ticket for every budget. The ranges around €22.75 and offers round-trip transfers to and from Athens. At around €36.90, you can book a , where you can enter the temple and learn about its cultural relevance from an expert tour guide.
  • Save big with Temple of Poseidon combo tours . On a one-time purchase, you can explore the Acropolis archaeological site along with the Temple of Poseidon. From round-trip transfers to expert guides, you can enjoy every facility with this purchase. 
  • You can customize your tour according to your preferences. Ideally, you should step out early so that you can reach the temple ruins by 1pm to 2pm. This will allow you to spend more time at the surrounding beaches and soak in the sunset views from the Temple of Poseidon. 
  • All round-trip transfers, private rentals, and public buses will drop you off around 150 meters away from the temple ruins. The pathway from the main entrance to the temple gate is steep and uneven. Wheelchair users cannot stroll their way up. It may also be challenging for visitors with mobility aids to navigate up the path. 

Which Temple of Poseidon ticket is best for you?

Temple of Poseidon & Cape Sounio Tickets

If you are on a budget

Go for: Sunset tour from Athens Duration: 5 hours Guide: Optional Entry access to the Temple of Poseidon: Not included

  • This budget-friendly tour is one of the best choices to enjoy sweeping views of the Aegean Sea on your way to Cape Sounion. 
  • Take advantage of the round-trip transfers to and from Athens and Cape Sounion for a hassle-free experience. You can choose between 3 pick-up points at checkout. 

Recommended tours:

Temple of Poseidon & Cape Sounio Tickets

If you love sunsets and photography

Go for: Sunset guided tours Duration: 5 hours Guide: English-speaking archaeologist guide Entry access to the Temple of Poseidon: Included

  • Witness gorgeous views of the Aegean Sea from Cape Sounion during golden hours. Capture the Doric columns of Poseidon’s Temple as the setting sun casts a soft glow on them. 
  • Enjoy round-trip transfers from Athens to Cape Sounion in luxurious, air-conditioned vehicles with free Wi-Fi on board.

Temple of Poseidon & Cape Sounio Tickets

If you are a history buff

Go for: Guided tours Duration: 4 hours Guide: Expert English-speaking guide Entry access to the Temple of Poseidon: Included

  • If you are interested in ancient Greek history, consider booking a guided tour to learn about the Temple of Poseidon and its significance in Greek mythology from an expert tour guide. 
  • Enter Poseidon’s Temple, marvel at its ancient Doric columns, and look through its intricate details. Also, enjoy scenic views of the Aegean Sea and coastal areas as you travel to Cape Sounion. 

Temple of Poseidon & Cape Sounio Tickets

If you want to explore beyond

Go for: HOHO tours and combo tickets Duration: 5 to 10 hours Guide: Expert English-speaking tour guide Entry access to the Temple of Poseidon: Not included

  • Explore the Acropolis archaeological site, the Parthenon, the Temple of Athena Nike, the Panathenaic Stadium, and Piraeus with combo tours. They help you save time and money compared to separate ticket prices. 
  • Combo: Hop-on-hop-off tour of Athens and Cape Sounion sunset tour
  • Combo: Acropolis, Acropolis Museum, Temple of Poseidon guided tour with entry tickets

Your visual guide to the Temple of Poseidon in Cape Sounion

Cape Sounion and the Temple of Poseidon

Inside the archaeological site of Cape Sounion

Temple of Poseidon & Cape Sounio Tickets

Temple of Poseidon

The ancient Greek sailors sought to appease the sea god Poseidon and built this magnificent temple around the 5th century BC. Unlike typical Doric columns, which tend to be thick and robust, the ones in Poseidon’s Temple are sleek and elegant. These columns were intentionally designed to be thinner than usual to allow more light. This made the temple look higher and was fitting for its dramatic clifftop location.

Cape Sounion and the Temple of Poseidon

Temple of Athena Sounias

The archaeological site of Cape Sounion houses another temple dedicated to the Greek goddess of wisdom and warfare. The co-existence of Athena’s shrine with Poseidon’s Temple confirms that ancient Greeks invoked different gods depending on their needs and circumstances. Most likely, sailors prayed to Poseidon for safe voyages and sought Athena’s blessings for favorable circumstances.

Cape Sounion and the Temple of Poseidon

Sounio Beach

Walk down to the beach after exploring the temple ruins so that you can unwind in peace as you soak in the glorious views of the Aegean Sea. You can also swim in the salt waters or sunbathe on the beach.

Cape Sounion and Temple of Poseidon

Ancient fortifications of Sounion

You will also find remnants of walls, towers, and gates on the archaeological site of Sounion. It is said that scores of ancient Greek military were stationed along this fortification to guard the food supplies from Eubea, Pontos, and the silver mines of Lavrion.

On your way to the Cape Sounion archaeological site

Along the Athens Riviera, you will be spoilt for choice with numerous beaches and iconic temples. Here are a few popular ones to dip your toes into before catching sunset views at Cape Sounion.

Cape Sounion and the Temple of Poseidon

Glyfada Beach

The Glyfada Beach is situated along the Athens Riviera and is one of the most popular beaches near Athens. From watersports, and beach bars, to restaurants, Glyfada offers it all. You can also simply relax in the waters while taking in sweeping views of the Saronic Gulf. 

Cape Sounion and the Temple of Poseidon

Lake Vouliagmeni

Besides beaches and shacks, you can also find natural wonders on your way to Cape Sounion, like Lake Vouliagmeni. This lake contains mineral-rich waters and does wonders for your stress levels. Relax by the water or swim in these waters before continuing your trip to Poseidon’s Temple. 

Cape Sounion and the Temple of Poseidon

Kavouri Beach

Surrounded by lush pine trees and rugged caves, Kavouri Beach offers a secluded escape near Cape Sounion. Kavouri’s small sea coves are perfect for snorkeling. If you are traveling with your partner, consider dropping by Kavouri Beach to steal away a few quiet moments for yourselves.

Cape Sounion and the Temple of Poseidon

Limanakia Beach

The Limanakia Beach is more of a series of rugged outcrops and small coves, ideal for sunbathing and snorkeling. The hidden inlets and dramatic cliff tops offer gorgeous views of the Aegean Sea. However, the pathway to Limanakia Beach is uneven and steep, so make sure to wear comfortable, grippy shoes when planning a visit.

Cape Sounion and the Temple of Poseidon

As you exit the city and drive south toward Poseidonos Avenue, you can spot some of the many Corinthian columns of the Temple of Zeus. As the name suggests, this temple was built to honor the King of the Olympian Gods. Its imposing structure, angular arches, and grand columns reflect the true grandeur of ancient Greek temples. 

Cape Sounion and the Temple of Poseidon

Archaeological site of Thorikos

The Thorikos archaeological site is one of the oldest mining areas in the world, situated near Cape Sounion. Their silver mines played an important role in facilitating trade and commerce in ancient Greece. In the present day, you can learn about the site’s history from these mines and the daily settlements of their workers. 

What to expect on a Cape Sounion tour from Athens?

A journey to the Temple of Poseidon in Cape Sounion is as much about the sunset views as the temple ruins. Here’s what you need to know about traveling from Athens to Poseidon’s Temple. 

  • Pick up and drop off: You will be picked up from your hotel or the Halandri metro station in Athens in air-conditioned vehicles. Check your ticket voucher for exact pick-up points and timings when booking tickets. It will take you around 1.5 to 2 hours to reach the temple ruins. 
  • Additional sites: The real charm of traveling to Cape Sounion is catching glimpses of the famous historic sites along the way. From the Acropolis archaeological site, Glyfada Beach, to Lake Vouliagmeni, and the Temple of Zeus, you can spot many other landmarks. 

Plan your visit to the Temple of Poseidon in Cape Sounion

Opening hours - Cape Sounion and Temple of Poseidon Tickets

  • Open: From 9:30am
  • Last admission: 20 to 30 minutes before sunset
  • Closed on: 1 January, 25 March, 7 May, Easter Sunday, 25, and 26 December
  • Good Friday: 12pm to 5pm
  • Holy Saturday: 9:30am to 3:30pm
  • Free admission inside the Temple: 6 March, 18 April, 18 May, European Heritage days (usually the last weekend of September), 28 October

Duration of visit: 20 to 40 minutes

Best time to visit: April to May and September to October, the spring and fall seasons, are the best months to visit Cape Sounion. You can avoid the scorching summer heat and spend more time exploring the outdoors without risking a heatstroke. Arrive at the site by afternoon and stay until after 5:30pm to catch stunning sunset views of the Aegean Sea.

Address: Cape Sounio, Sounion 195000, Greece Find on maps

  • By round-trip transfers: Choose round-trip transfers from Athens. It helps you save time and money, and you can comfortably be seated on luxurious, air-conditioned vehicles, as you make your way to the site.
  • By car: The city is around 77km away from Cape Sounion. If you drive along Highway A6 , you have to pay toll fees. However, if you opt to ride through the coastal stretch of Athinon Souniou Road , it will take you an extra 15 to 20 minutes to reach the site, but you can skip the toll charges.  You can also spot several iconic landmarks and historic sites along the way. 
  • By bus: Another budget-friendly way to reach the Temple of Poseidon is to board KTEL buses . These public buses run from Athens to Cape Sounion once or twice a day. You can board them from Mavromateon Terminal , one of the KTEL bus stops on the western side of Pedion Areos Park . To reach this terminal, get down at the Victoria station on the M1 line. 

Temple of Poseidon & Cape Sounio Tickets

  • Main public entrance: The main entrance to the Temple of Poseidon is situated at the base of the cliff. There is an information center and washroom stalls nearby. There are no specific demarcation or information signs around this spot. You will find cars parked along the way, until the entry gate. 
  • Pathway to the temple ruins: The road leading from the entrance to the temple ruins is steep and uneven. While the pathway is well-maintained and equipped with railings and resting spots on the way up, it may be challenging for elderly tourists and visitors with limited mobility to navigate. The path winds its way through the Sounion archaeological site and offers great views of the Aegean Sea and surrounding beaches. You can also spot the smaller Temple of Athena Sounios on the way up. 

Temple of Poseidon & Cape Sounio Tickets

  • Cafe: There is a cafe at the bottom of the hill, a short walk from the parking area and information center. They offer light pita sandwiches, Koulouri, a type of sesame bread ring, Loukoumades, Greek donuts with honey, and a variety of nuts, and dried fruits. While not overlooking the Aegean Sea, this cafe offers scenic views of the surrounding areas. 
  • Visitor center: There is an information center at the bottom of the hill. You can find information about comfortable transfers, bus routes, opening hours, and other details there. 
  • Toilets: You can use washroom facilities at the information center or the cafe, at the bottom of the hill. 

Temple of Poseidon & Cape Sounio Tickets

If you book Cape Sounion sunset tours, they often include an expert tour guide, who will also help visitors with limited mobility board the car. However, the pathway from the main entrance to the temple gate is not wheelchair-friendly. The road is steep and uneven. We recommend you bring along a friend, family member, or caregiver to assist you along the way. 

  • Steps and ramps: Visitors who have walking aids, but are not completely wheelchair-bound, may attempt to walk up the hill. There are well-maintained steps and ramps in certain areas. Also, you do not have to walk all the way up. You can spot heritage sites and see the crashing waves only a few steps in. 
  • Handrails and resting spots: The pathway has handrails and armrests so that you do not lose balance. There are well-positioned tree barks and benches at frequent intervals. You can rest your feet, have a snack, and begin your hike again. 

Temple of Poseidon & Cape Sounio - Rules

  • Stay on marked paths: Cape Sounion has designated pathways to protect certain areas of the temple ruins and to ensure the safety of its visitors. Be careful so as not to stray from these marked pathways. 
  • Leave no trace: You can munch on light snacks as you make your way up the hill, however, do not throw any wrappers or plastic bottles on the road. 
  • Maintain photography etiquette: While it is tempting to take selfies at every beautiful corner of Cape Sounion, be mindful of other visitors. Do not take pictures in restricted areas. Avoid bringing selfie sticks, heavy camera gear, and drones to the tour. 

Temple of Poseidon & Cape Sounio - Visitor tips

  • Arrive early at the bus stop: If you are planning to travel to Cape Sounion from Athens on public KTEL buses, keep a few minutes on hand. These buses usually depart at 10am from Mavromateon Terminal, however, in some cases, they may also leave the stop around 9:30am. If you aren’t sure where to wait for the bus, please ask any passerby. 
  • Use restrooms before trekking up: It will take you around an hour to walk up from the main entrance to the temple ruins. It is best to grab a snack or use the restroom before making your way up, as the information center and cafe close around 5:30pm. 
  • Pack light: The Temple of Poseidon does not have a cloakroom or storage facility on site. Avoid bringing suitcases or backpacks on your tour. 
  • Bring a jacket: The temple ruins on the top of the hill tend to get windy around sunset hours. Bring a scarf or jacket, especially if you are traveling around the fall and winter months. 
  • Check the weather forecast: One of the major highlights of traveling to Cape Sounion is the scenic views of the Aegean Sea. Avoid traveling on foggy days as your visibility may be obstructed. 

Eat - Cape Sounion and Temple of Poseidon Tickets

  • Naos Cafe (280m away): This cafe, situated at the bottom of the Temple of Poseidon’s hill offers a selection of light snacks and beverages. Expect filling pita sandwiches, souvlaki, a type of grilled meat skewers, and fresh Greek salads, drizzled with olive oil. 
  • The Veranda Restaurant (2.1km away): This restaurant offers a delectable range of seafood. Look out for their shrimp saganaki, baked with cheese and tomatoes, and their moussaka, a layered eggplant dish. 
  • Ilias (2km away): The Ilias is a family-run tavern, situated a few blocks from the temple ruins. Try their classic seafood preparations, made with authentic Greek flavors. Their spicy calamari (fried squid), dolmades (stuffed grape leaves), and pastitsio (a layered pasta dish with meat and bechamel sauce) are some house favorites. 
  • Cape Sounio Restaurant (4.2km away): This upscale restaurant offers a fine dining experience with gorgeous views of the Aegean Sea. Their locally caught fish, grilled or baked, with a lemon-olive oil dressing is a must-try. 
  • Akrogiali (17km away): This establishment is around a 20-minute drive away from the Temple of Poseidon. Try their grilled sardines, topped with olive oil and lemon, and Horiatiki, a fresh salad prepared with tomatoes, peppers, olives, and a generous feta dressing. 

Nearby - Cape Sounion and Temple of Poseidon Tickets

  • Sounio Beach (adjacent to the Temple of Poseidon): This beach is nestled on a bay and offers spectacular views of the Temple of Poseidon and the Aegean Sea. The gentle waves and clear waters make it a perfect spot for families to swim and sunbathe. 
  • Temple of Athena Sounio (200 m away): Built during the 5th century BC, this smaller temple is located on the same site as the Temple of Poseidon. Although in ruins, this temple offers visitors a closer look at the polytheistic nature of worship in ancient Greece. 
  • Archaeological Museum of Lavrion (10 km away): If you are interested in learning more about the history of the Cape Sounion region and why its fortifications were built, head to the Lavrion Archaeological Museum. They showcase artifacts about Sounion’s silver mines and how they functioned in ancient Greece.
  • Lake Vouliagmeni (22 km away): Situated in the coastal town of Vouliagmeni, this lake offers a serene escape from the hustle-bustle of city life. The lake is fed by natural springs and its water is rumored to contain healing properties. 

Frequently asked questions about Temple of Poseidon tickets

It is best to book Temple of Poseidon tickets and Cape Sounion tours online to ensure a hassle-free experience. Pre-booking tickets helps you save time and money. You can skip ahead of long ticketing lines at the venue and directly enter the temple ruins.

Yes, if you purchase Temple of Poseidon tickets online, you can enjoy reduced prices and combo offers. On a one-time purchase, you can explore many historic landmarks in and around Cape Sounion. With online ticket bookings, you can also enjoy priority access to the temple ruins.

You can travel to Cape Sounion from Athens either by car or public bus. The KETL public buses run once or twice a day from Mavreomateon Terminal on the western side of Pedion Areos Park. Arrive by 9:30am to ensure you do not miss the bus. It will take you around an hour and a half to reach Sounion via car. The Highway A6 route has tolls, however, it is shorter than the coastal Athinon Souniou Road.

Depending on the kind of Temple of Poseidon tickets you book, you can enjoy round-trip transfers and priority access to the temple ruins. Travel in private, air-conditioned vehicles with an expert tour guide from Athens to Sounion. You will also receive free Wi-Fi and basic travel insurance on board. 

Athens is almost 77km away from Cape Sounion. You need around 3 to 4 hours to make the most of your visit to Cape Sounion. We recommend you pick the longer, scenic, Athinion Souniou Road to spot iconic landmarks along the way. Upon arrival, spend around 30 minutes to an hour exploring the temple ruins and head to the nearby Sounio Beach. Swim in the waters and sunbathe. Head back to the temple ruins around sunset and soak in the golden hour views of the Aegean Sea.

March to May and April to September are the best months to travel to Cape Sounion. The weather is pleasant and crowds are manageable. Arrive by 5pm at the archaeological site of Cape Sounion to enjoy sunset views of the Aegean Sea. Check the weather forecast before planning a visit. Rain and foggy weather will disrupt your visibility. 

The Temple of Poseidon is open from 9:30am till the sun sets on most days throughout the year. Usually, it closes by 5:30pm. During summer, the site is open until around 6:30pm to 7pm as the days are longer. The temple is closed on 1st January, 25th March, Easter Sunday, 7th May, 25th, and 26th December. On Good Friday, the temple is open from 12pm to 5pm. And, on Holy Saturday, the temple is open from 9:30am to 5:30pm.

The Temple of Poseidon is perched atop Cape Sounion, around 60 meters above the Aegean Sea. The temple’s location marks the southernmost tip of the Athenian territory. The dramatic interplay of the Aegean Sea and its surrounding rugged cliffs makes for gorgeous views from the temple ruins of Cape Sounion.

Temple of Poseidon Tour

Temple Of Poseidon Sunset Tour

Overview of temple of poseidon.

The Temple of Poseidon Sunset Tour involves exploring the temple that is housed within a fort that guards Attica's coastline and is located at the southernmost point of the Sounio Peninsula. It was constructed in the middle of the 5th century BC in honor of Poseidon, the sea god, and is entirely made of white marble. The shades of blues, reds, and oranges that envelop the temple as the sun sinks into the water will mesmerize you. One of many who succumbed to its charm was the British poet Lord Byron, who wrote poetry in praise of it and had his name inscribed on one of its recognisable marble pillars.

To get to the historic site of Temple of Poseidon Sunset Tour, take the roads along the southern Attica coastline. Discover the temple's ruins and enjoy expansive views of the Aegean before witnessing a stunning sunset. Enjoy a relaxing 50-minute trip in an air-conditioned minivan along Attica's coastal roads as you take in the picture-perfect beaches and sleepy towns as you pass. After arriving at Cape Sounion, proceed to the old Poseidon temple, which is devoted to the Greek deity of the sea, and have a tour of this important historical site. Before taking some time to unwind and enjoy the expansive views of the Aegean, discover the folklore associated with the temple.

Book Temple Of Poseidon Sunset Tour Online

Yes, you can schedule a Temple of Poseidon Sunset tour in advance online. By booking in advance, you can purchase tickets while relaxing in your home or hotel room. This guarantees availability and makes it easier for you to browse through several combo packages and select the one with the best deal. One of the biggest benefits of booking online is that you may schedule your trip in advance and avoid the lengthy lines.

With the Temple of Poseidon Sunset tour discover the history hidden behind each curve as you travel the Athens Riviera with a guide. On Cape Sounion, watch the sunset over the Aegean Sea and visit the Poseidon Temple. From the Halandri Metro station in Athens, take a bus to Cape Sounion to arrive in time for the captivating sunset.

Book Now - Acropolis Of Athens Tickets Online

Highlights For Temple Of Poseidon Sunset Tour

Highlights For Temple Of Poseidon Sunset Tour

  • Visit the magnificent Temple of Poseidon on Cape Sounion in the evening and take in the Aegean Sea sunset.
  • From the Halandri Metro station in Athens, take a bus to Cape Sounion to arrive in time for the captivating sunset.
  • Your archaeologist guide will teach you about Sounion's recent history and current situation.
  • Discover the Athens Riviera while getting a taste of everyday life there.
  • Visit the Poseidon Temple that rises from Cape Sounion's edge.
  • View the breathtaking sweeping panorama from Sounion's southeast peak.
  • Experience the stunning Aegean Sea sunset.
  • Check out the great authors' signatures that they left behind on the temple's marble, including the Romantic poet Lord Byron.
  • Make time to visit Lake Vouliagmeni, a special natural gem on the Athenian Riviera.

Also Visit - Hop-On Hop-Off Bus & Cape Sounion Sunset Tour

FAQs for Temple Of Poseidon Sunset Tour

What is the temple of poseidon.

The Temple of Poseidon is an archaeological site on Cape Sounion which was built in the 5th century BC in honor of the Greek mythological God of the Sea. The columns, which were constructed of white marble, continue to exist as a memorial of the Athenian Empire.

Also Checkout - Parthenon Greece

Why is the Temple of Poseidon so famous?

The renowned Temple of Poseidon as its name suggests is dedicated to Poseidon, the Olympian God of the Sea. It was built on Aegina Island during the Golden Age of Pericles. Along with the renowned sites of Parthenon and the temple of Aphaia, it makes up the Sacred Triangle of antiquity. It offers breathtaking views of the Aegean Sea due to its cliffside location. One of the most unforgettable experiences is watching the sunset from the temple which paints a beautiful picture over the Aegean Sea.

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How much time is needed for the Temple of Poseidon tour?

To visit the breathtaking Temple of Poseidon Sunset tour on Cape Sounion in the dusk, a five-hour tour is necessary. In these five hours, you may take in the Athenian culture, learn about the Athens Riviera, and watch the sunset over the Aegean Sea.

Do Checkout: Temple of Zeus Tickets

Can we book a Temple of Poseidon tour online?

Yes, you can book a Temple of Poseidon Sunset tour online, in advance. Booking in advance helps you book tickets from the comfort of your homes or hotel rooms. You can also book tickets on the go while you are traveling. This ensures availability and also helps you browse through various combo packages so that you can choose the best price.

Do Checkout: Temple of Olympian Zeus

What are the top places to visit near the Temple of Poseidon?

When you set out on the Temple of Poseidon Sunset tour make sure you tick off other Athens landmarks near the temple. These include the New Acropolis Museum – a place that houses many masterpieces and the Temple of Zeus, the Olympian God. You should also explore the main square of Athens, Hadrian’s Arch, Tomb of Unknown Soldier, The Academy of Athens and Maximos Mansion. To get away from the hustle bustle and to blow off steam, spend some time at the Sounio Beach, Kape Beach and Paralia Christea.

What are the best tours that we can book while visiting Temple Of Poseidon?

Here is the list of best Temple Of Poseidon tours:

  • Temple Of Poseidon Private Tour
  • Cape Sounion & Temple Of Poseidon Sunset Tour
  • Hop-On Hop-Off Bus & Cape Sounion Sunset Tour
  • Cape Sounio & Poseidon Temple Trip With Audio Guide
  • Cape Sounion & Temple Of Poseidon Half Day Tour
  • Cape Sounion & Temple Of Poseidon Afternoon English Tour
  • Full Moon Tour Of Temple Of Poseidon

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Savvino-storozhevsky monastery and museum.

Savvino-Storozhevsky Monastery and Museum

Zvenigorod's most famous sight is the Savvino-Storozhevsky Monastery, which was founded in 1398 by the monk Savva from the Troitse-Sergieva Lavra, at the invitation and with the support of Prince Yury Dmitrievich of Zvenigorod. Savva was later canonised as St Sabbas (Savva) of Storozhev. The monastery late flourished under the reign of Tsar Alexis, who chose the monastery as his family church and often went on pilgrimage there and made lots of donations to it. Most of the monastery’s buildings date from this time. The monastery is heavily fortified with thick walls and six towers, the most impressive of which is the Krasny Tower which also serves as the eastern entrance. The monastery was closed in 1918 and only reopened in 1995. In 1998 Patriarch Alexius II took part in a service to return the relics of St Sabbas to the monastery. Today the monastery has the status of a stauropegic monastery, which is second in status to a lavra. In addition to being a working monastery, it also holds the Zvenigorod Historical, Architectural and Art Museum.

Belfry and Neighbouring Churches

poseidon temple tour

Located near the main entrance is the monastery's belfry which is perhaps the calling card of the monastery due to its uniqueness. It was built in the 1650s and the St Sergius of Radonezh’s Church was opened on the middle tier in the mid-17th century, although it was originally dedicated to the Trinity. The belfry's 35-tonne Great Bladgovestny Bell fell in 1941 and was only restored and returned in 2003. Attached to the belfry is a large refectory and the Transfiguration Church, both of which were built on the orders of Tsar Alexis in the 1650s.  

poseidon temple tour

To the left of the belfry is another, smaller, refectory which is attached to the Trinity Gate-Church, which was also constructed in the 1650s on the orders of Tsar Alexis who made it his own family church. The church is elaborately decorated with colourful trims and underneath the archway is a beautiful 19th century fresco.

Nativity of Virgin Mary Cathedral

poseidon temple tour

The Nativity of Virgin Mary Cathedral is the oldest building in the monastery and among the oldest buildings in the Moscow Region. It was built between 1404 and 1405 during the lifetime of St Sabbas and using the funds of Prince Yury of Zvenigorod. The white-stone cathedral is a standard four-pillar design with a single golden dome. After the death of St Sabbas he was interred in the cathedral and a new altar dedicated to him was added.

poseidon temple tour

Under the reign of Tsar Alexis the cathedral was decorated with frescoes by Stepan Ryazanets, some of which remain today. Tsar Alexis also presented the cathedral with a five-tier iconostasis, the top row of icons have been preserved.

Tsaritsa's Chambers

poseidon temple tour

The Nativity of Virgin Mary Cathedral is located between the Tsaritsa's Chambers of the left and the Palace of Tsar Alexis on the right. The Tsaritsa's Chambers were built in the mid-17th century for the wife of Tsar Alexey - Tsaritsa Maria Ilinichna Miloskavskaya. The design of the building is influenced by the ancient Russian architectural style. Is prettier than the Tsar's chambers opposite, being red in colour with elaborately decorated window frames and entrance.

poseidon temple tour

At present the Tsaritsa's Chambers houses the Zvenigorod Historical, Architectural and Art Museum. Among its displays is an accurate recreation of the interior of a noble lady's chambers including furniture, decorations and a decorated tiled oven, and an exhibition on the history of Zvenigorod and the monastery.

Palace of Tsar Alexis

poseidon temple tour

The Palace of Tsar Alexis was built in the 1650s and is now one of the best surviving examples of non-religious architecture of that era. It was built especially for Tsar Alexis who often visited the monastery on religious pilgrimages. Its most striking feature is its pretty row of nine chimney spouts which resemble towers.

poseidon temple tour

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The Unique Burial of a Child of Early Scythian Time at the Cemetery of Saryg-Bulun (Tuva)

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Pages:  379-406

In 1988, the Tuvan Archaeological Expedition (led by M. E. Kilunovskaya and V. A. Semenov) discovered a unique burial of the early Iron Age at Saryg-Bulun in Central Tuva. There are two burial mounds of the Aldy-Bel culture dated by 7th century BC. Within the barrows, which adjoined one another, forming a figure-of-eight, there were discovered 7 burials, from which a representative collection of artifacts was recovered. Burial 5 was the most unique, it was found in a coffin made of a larch trunk, with a tightly closed lid. Due to the preservative properties of larch and lack of air access, the coffin contained a well-preserved mummy of a child with an accompanying set of grave goods. The interred individual retained the skin on his face and had a leather headdress painted with red pigment and a coat, sewn from jerboa fur. The coat was belted with a leather belt with bronze ornaments and buckles. Besides that, a leather quiver with arrows with the shafts decorated with painted ornaments, fully preserved battle pick and a bow were buried in the coffin. Unexpectedly, the full-genomic analysis, showed that the individual was female. This fact opens a new aspect in the study of the social history of the Scythian society and perhaps brings us back to the myth of the Amazons, discussed by Herodotus. Of course, this discovery is unique in its preservation for the Scythian culture of Tuva and requires careful study and conservation.

Keywords: Tuva, Early Iron Age, early Scythian period, Aldy-Bel culture, barrow, burial in the coffin, mummy, full genome sequencing, aDNA

Information about authors: Marina Kilunovskaya (Saint Petersburg, Russian Federation). Candidate of Historical Sciences. Institute for the History of Material Culture of the Russian Academy of Sciences. Dvortsovaya Emb., 18, Saint Petersburg, 191186, Russian Federation E-mail: [email protected] Vladimir Semenov (Saint Petersburg, Russian Federation). Candidate of Historical Sciences. Institute for the History of Material Culture of the Russian Academy of Sciences. Dvortsovaya Emb., 18, Saint Petersburg, 191186, Russian Federation E-mail: [email protected] Varvara Busova  (Moscow, Russian Federation).  (Saint Petersburg, Russian Federation). Institute for the History of Material Culture of the Russian Academy of Sciences.  Dvortsovaya Emb., 18, Saint Petersburg, 191186, Russian Federation E-mail:  [email protected] Kharis Mustafin  (Moscow, Russian Federation). Candidate of Technical Sciences. Moscow Institute of Physics and Technology.  Institutsky Lane, 9, Dolgoprudny, 141701, Moscow Oblast, Russian Federation E-mail:  [email protected] Irina Alborova  (Moscow, Russian Federation). Candidate of Biological Sciences. Moscow Institute of Physics and Technology.  Institutsky Lane, 9, Dolgoprudny, 141701, Moscow Oblast, Russian Federation E-mail:  [email protected] Alina Matzvai  (Moscow, Russian Federation). Moscow Institute of Physics and Technology.  Institutsky Lane, 9, Dolgoprudny, 141701, Moscow Oblast, Russian Federation E-mail:  [email protected]

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